Struggling to find their purpose on the Betterman farm, the Croods see an opportunity to prove their worth by harvesting Phil’s perilous (and sticky) bounceberries.
After a crowverine gliding mishap, Dawn loses her confidence and tells Eep she's taking a break from adventures. Eep convinces Dawn to join her on one last adventure – climbing Grrr Mountain.
Ugga has a secret - she's taking Eep to "flop with the bulrusses" - a fun and dangerous pastime Ugga enjoyed as a teen. Ugga's excitement fades when Hope invites herself and Dawn along.
Bored of the viewing options on window, Thunk realizes what he’s been missing – real life drama! But when Thunk starts creating drama around the farm, the others set out to teach him a lesson.
Guy is always surprising Eep with romantic gestures and Eep wants to return the favor. But when Guy proves impossible to surprise, Eep has to step up her game.
The Crood and Betterman parents have been struggling to get along – they’re so different! The two couple realize they need some time apart and decide to get away from the farm and each other.
While the parents are away, the kids are in charge of the tree house, but when the punch monkeys invade the house, the kids find themselves out of their element.
When Eep and Dawn find a stranded egg, they eagerly take it in as their new pet, but when they learn the egg is not what it seems, they have to return it before it becomes dangerous.
When Phil invents a new game called ball-in-cup, everyone except Hope can't stop playing; just as Hope thinks she's rid the farm of the game, she finds herself addicted.
When a freak lightning strike turns Thunk into a genius, Phil is thrilled to have someone like-minded on the farm -- until the residents start asking Thunk for help with problems instead of him.
Tired of the farm's feud with the neighbouring punch monkeys, Ugga arranges a peacemaking summit where the Bettermans and Croods will stay in the punch monkey village to learn how to get along.
Determined for Grug and Guy to bond, Eep plans a hunt for the three of them. But when Eep sneakily makes it appear that Guy saved Grug from a charging bearacuda, the two believe Guy is a master hunter, and head to hunt a dangerous scorpulion.
When Grug saves Hope's life during a dangerous mountainside tartichoke harvest, rather than feeling grateful, Hope is desperate to save Grug's life in order to not be indebted to him.
Determined to show Hope that she's just as good a cook, Ugga decides to cook a meal that will wow the farm residents. But when Ugga realizes she's out of her element, she turns to the punch monkeys for help.
When Hope allows Dawn to borrow her favorite bracelet, Dawn is determined to prove she is responsible. But when an accident leads to the bracelet being stuck in an Eegle's nest, Dawn, Eep, and Guy must retrieve the bracelet before Hope finds out.
Gran announces it's time for a Thunder Sister retreat at Camp Thunder to strengthen their team building skills, but Gran takes it too far when she puts the sisters through a dangerous trial.
Each kid believes that they have the best pet on the farm, and hold a contest to prove it. Meanwhile, Phil's attempts to find a pet to enter proves dangerous, as he turns to wild animals.
After a back-breaking day of farm chores, all Grug and Phil want is a slice of pie. But when there is only one slice left, both Grug and Phil argue that they deserve it more. To prove who works harder, Grug and Phil switch chores on the farm for a day.
Dawn is bummed when an injury prevents her from joining the Croods and Bettermans to a trip to Fizzy Falls, but when Thunk recruits her on his quest for an ancient stash of legendary snacks, Dawn can't resist joining the treasure hunt.
Believing he doesn't fit in on the farm, Chunky (aided by the other pets) leaves home to follow the distant call of another death cat. The kids set off to bring him back home. Note: Gran narrates the story.
Grug accidentally reveals a subterranean cave beneath the treehouse. When Phil can't convince Hope, Grug, and Ugga that the treehouse is more impressive than the new cave, he sets out to destroy the cave.
The kids take their "Dare Game" too far, causing Grug, Ugga, Phil and Hope to forbid them from playing it. But when the adults also get hooked on the game, the kids must convince them to stop before someone gets hurt.
Thunk goes missing after the Thunder Sisters dismiss his help during a battle. The Thunder Sisters, and Grug, Phil, and Guy's newly formed "Thunder Sifters" search on and off the farm for Thunk.
The Croods, Bettermans, and Guy compete in the Throw-Go-Bowl - an annual Throw-go-nut championship game against the punch monkeys. Grug's desire to win above all else threatens to push away his teammates.
Phil enlists the Croods to help him retrieve a rare fruit before some wild animals can. But when Grug doesn't trust Phil's methods of getting it back to the farm, the Croods and Bettermans compete with each other more than with the animals.
When the parents find their precious outhouse destroyed, they perform a "trouble trial" to find the culprit. To prove their innocence, each kid provides their own explanation for the outhouse's destruction.
After a big argument between the Croods and Bettermans, the parents decide to separate the two families. Not wanting to split up, the kids try to convince the parents to make up before the Croods move off the farm entirely.
Hwam returns with news of a utopian cave community, accidentally reigniting the argument between the Croods and Bettermans. As the Croods journey to Caveana and the Bettermans adjust to life without them, the parents deny how much they miss each other.