Mick arrives at Josef’s office building, which is on fire. Nothing is left intact. No survivors. Someone who knew Josef was a vampire attacked him with one of the two known ways to kill a vampire. Mick and Tom, Josef’s head of security, launch in investigation into the attack. Beth does her own investigating and interviews several people from Josef’s past, none of whom have anything nice to say. The twin investigations lead Mick and Beth to New York, and yet another surprise from Josef’s past.
Josh is working on the biggest case of his career, a conviction against a member of the MS-13 gang, possibly the most dangerous gang in the world. In an attempt to force Josh to drop the case, the gang threatens Beth, but Josh refuses to fold and enlists Mick’s help in protecting her. Unfortunately, Josh is left unguarded and is kidnapped with devastating results. Beth discovers an unwanted blood connection.
While putting Josh's affairs in order, Beth discovers evidence leading her to believe Josh was cheating on her. When she investigates, however, Beth learns that Josh was about to propose to her and that the woman she thought he was cheating with was actually a consultant he hired for a confidential case - a case that turns out to involve Mick. Coraline's disappearance from the hospital, with the assistance of an old friend, leads Mick to the discovery of a temporary 'cure' for vampirism and a surprising link to the French revolution. Meanwhile, a startling family connection of Coraline's is revealed.
Benjamin Talbot, a new A.D.A., investigates the murder of Beth's boss Maureen. Beth and Mick pursue their own investigation, Benjamin warns them not to spoil any evidence that could lead to the culprit. Mick provides some good advice and some surprising empathy to a troubled teenager Bonnie with secrets of her own.
A human is killed by a vampire in the heat of passion, causing Mick and Beth to face the ultimate consequences of a vampire-human relationship. The investigation of the murder puts Mick and Josef's secret identities in jeopardy, forcing them to join forces with the other vampires in Los Angeles to keep their secret safe.