土門は不正な手段を使ってヤマト配属を叶えていた。第65護衛隊、出航の日。新たなるクルーの手で、ヤマトが動き出す。機関室には、あの徳川機関長の息子、徳川太助。そして舵を握るのは、土門。ヤマト艦長となった古代進は、彼に艦を託すことでその覚悟を問う。
Domon had obtained his assignment to Yamato through illegal means. The day of the 65th convoy's departure. The Yamato is set in motion by a new crew. In the engine room is Tokugawa Taisuke, the son of the chief engineer Tokugawa. And at the helm is Domon. Susumu Kodai, now Yamato's captain, questions his resolve by entrusting the ship to him.