Sonny Munroe enters the set, not knowing that she is in for a difficult time with Tawni Hart. When Sonny changes Tawni's "Queen Bee" sketch, Tawni becomes jealous; the situation does not improve when Sonny and Tawni have to share a dressing room. Sonny's mother, Connie (Nancy McKeon) advises her to talk to Tawni. Things get worse when Sonny tries to help Tawni, as she accidentally shreds Tawni's favorite stuffed animal. Then, when Tawni talks to Marshall (Michael Kostroff), Sonny decides she doesn't care what Tawni thinks anymore and makes the One Bad Bee sketch to get back at her.
Sonny feels left out when her best friend from Wisconsin comes to visit and instantly hits it off with Tawni. Tension reaches the boiling point when Tawni and Lucy go to Chad's birthday party... without inviting Sonny. Meanwhile, Nico and Grady plot revenge against a security guard who has been helping himself to their pizza.
The "So Random!" cast attempt to prevent their prop house from being taken away and used as a meditation room for the "MacKenzie Falls" cast. The number one show gets any room in the building according to the contract. After remembering all the good memories about sketches they have done (like Dolphin Boy and the Check It Out Girls), they hand-cuff themselves to the prop house. After two hours of being hand-cuffed they decide to fight back. They chase away the cast of "MacKenzie Falls" by cutting a hole in the ceiling and annoying them by throwing objects on their heads. Chad says he won't be driven away, but he starts meditating in the photo booth, he runs away because of a rat. The So Random cast get their prop house back. Chad comes back and brings gifts to everyone in order to say he is sorry and didn't know how much the prop house meant to them. He gives Tawni a Bedazzled lipstick case, Zora a night light for the sarcophagus, lets Nico and Grady ride the forklift, and lastly gives an autograph picture of himself to Sonny that says "To my biggest fan, I'm sorry, TV's Chad Dylan Cooper." Then Sonny says she is touched. Then, Nico and Grady bursts in while riding the forklift.
Sonny tries to help Tawni have a good date with an intern Hayden. Sonny begs Chad for his Lakers tickets. But when she and Hayden are shown on the kiss cam they are forced to kiss and Tawni sees them. Tawni gets mad and hides in her dressing room. Meanwhile, Chad gets jealous because he sees the kiss in Tween Weekly Magazine. Sonny makes another date and tells Hayden that Tawni is going to cook for him. But when Sonny learns that Tawni can't cook, Sonny offers to have the date at her place. After Sonny ruins the food, she tries to call Nico and Grady to bring food but she drops her phone in the sink. Tawni tries to help her but turns on the garbage disposal. Chad thinks that Sonny is in trouble and later breaks down her door. Meanwhile, Grady and Nico try to put their influence on the name of a sandwich but whoever ate the sandwich ends up puking.
Chad is the new guest star on So Random, and he and Sonny have to do a sketch together where they will kiss, much to her dismay. Chad then says when Sonny looks into his eyes she will fall in love with him. Sonny takes this seriously, and, worried she will fall in love with Chad tries everything she can to get out of it. In the end, they do the sketch and just as their about to kiss a pig flies into Sonny's hands, so she let's the pig kiss Chad. Meanwhile, Nico and Grady get control of Marshall's office for the day, in exchange for Chad's So Random appearance.
Sonny, who once was a Blossom Scout herself but got her sash taken away, helps Zora break a record of selling the most cookies but the studio owner's daughter joins the Blossom scouts just to see Zora lose and bribes Chad to help her. Tawni takes a bet from Sonny that she can't stay away from looking at herself in the mirror. Nico and Grady create their own scent to get girls.
When Sonny and Tawni are guests on "Gotcha with Gilroy Smith," Gilroy tries to get Tawni and Sonny to confess some embarrassing moments on "So Random!". Sonny and Tawni make a pact backstage that they will have each others back but they realize that there is a hidden camera so Gilroy knows about their pact. Tawni breaks the pact and Sonny starts to think about Chad Dylan Cooper, but Gilroy knows that is a face of someone who has a boy crush. Gilroy brings Chad out (Chad thinks he's there because of a book he read) and shows a clip of him and Sonny sharing a rather romantic moment from Episode 2. They both go backstage and they don't realize there's another hidden camera and everyone is watching them. Sonny and Chad are about to tell each other what their feelings are for one another, but Tawni, feeling she owes Sonny, grabs the remote and turns off the camera so Gilroy and the studio audience can't see what happens. Gilroy turns the camera back on and sees Sonny saying "That was so much easier to say than I thought it would be," and then walking away with Chad. Gilroy becomes mad because he doesn't know what Sonny and Chad said to each other, although it is thought that they told each other that they like each other, based on what they were saying before and after Tawni turned the camera off.
Sonny and Tawni are invited on an exchange program to Glendovia because of the success of their Check-it-Out Girls sketch, but refuse to go together due to an unknown argument. When Tawni goes she is shocked to discover that it is a cultural wasteland and that the Glendovians think the characters are real check out girls. Meanwhile, an exchange student impresses a girl Nico and Grady like, and they both think it is because of his mustache. They then wear a mustache trying to impress her, but they get themselves stuck in their own mustaches because of the glue.
Feeling that Gassie doesn’t get the affection that he deserves, Sonny takes it upon herself to show the dog love, love that it doesn’t get from its trainer. Sonny then finds out that giving the dog affection effects Gassie’s mood to flatulate on command when filming the “Gassie” sketches as well jeopardizing the feature film “Gassie & We”. Sonny then gives Gassie too many dog treats that may have killed him. In the end it turns out Gassie was in shock from too many meatball treats and gets the love he deserves, also putting Chad into shock by pretending to be dead. Meanwhile, Nico and Grady borrow money from Dakota Condor to invest in fart noise lunch boxes for “Gassie & We”.
Sonny and her fellow cast members are forced to attend a public high school after they are all fired from the show because of their complaints about a new addition to the cast. Can Tawni, Nico, Grady, and Zora survive a normal high school, even though it's not the "musical" they thought it would be?
When Grace, an elderly woman, wins the “Be So Random! for a Day” contest, the “So Random!” cast ends up planning an “old geezer” sketch that not only offends Grace and her entire retirement home, but all elderly people. Grace and her friends turn the tables and play a practical joke on the cast. Meanwhile, Chad attempts to help a group of kids with their fear of clowns and ends up looking like one himself.
Someone on the set of So Random! and Mackenzie Falls starts to create rumors that Sonny is a thief when she temporarily leaves for her Homecoming in Wisconsin. This ruins her reputation, and she is asked to leave So Random!. Tawni supports Sonny as they drive back to Wisconsin, unable to clear her name. Meanwhile, Chad, Nico and Grady find out the truth back in Hollywood.
Sonny finds out that the studio head, Mr. Condor, has brought the show “Pauly the Polar Bear’s Playroom,” to shoot on the lot. The show happens to be Sonny’s favorite childhood show, so she introduces her self to her beloved Pauly. However, the guy who plays Pauly quits and makes Sonny wear the costume and pretend to be Pauly. Trapped in the Pauly costume meeting kids, she must convince the man who played Pauly to continue his career for the sake of his inspired fans. Meanwhile, it is Sonny and Chad’s seven week aniversary as a couple, but Sonny can’t make the date due to being Pauly. Nico and Grady take advantage and tell Chad that Sony may be losing interest, only so they can get him to join their video gamer’s group.