A young girl named Elizabeth appears in Gallery Fake, claiming to be Fujita's daughter. Unsure about the story, Fujita takes Elizabeth back to his apartment, but that night, several thugs break into his apartment and abduct her. Using a key she dropped, Fujita collects packet left by Elizabeth which includes the diary of Antonio de Beatis in a railway locker. He and Sara travel to Florence, Italy, to visit d'Avalos. He is an obsessive collector of copies of Leonardo da Vinci's painting known as the Mona Lisa. He also claims to be the descendant of the woman in the painting, Constanza d’Avalos, which helps his political ambitions. Fujita enlists Carlos' help and he provides some explosives. The diary suggests the painting is of another woman, Lisa Gherardini, the wife of Francesco del Giocondo. She is the ancestor of Elizabeth's mother, Flora del Anton Maria del Giocondo. Meanwhile d’Avalos has Elizabeth trapped in a tower on his estate.
三田村に連れられて、突然ギャラリーフェイクに現れた少女・エリザベータ。彼女はなんと、フジタの娘だという。仕方なくエリザベータを自分のアパートに連れて帰るフジタ。だがその夜、アパートに数人の暴漢が乱入。エリザベータを連れ去ってしまう。
エリザベータが残したある包みを手にしたフジタは、彼女をさらったダヴァロスを訪ねるべく、イタリア・フィレンツェへと向かう。フィレンツェの名士・ダヴァロスは、世界中の「モナリザ」を集めていたが…。