Surly, a mischievous squirrel, must plan a heist to get into his town's biggest nut shop in order to help his pals in the park gather food to survive the winter. Together with his sidekick, Buddy, he will assemble a ragtag crew to help him get inside — and take them on a fun-filled adventure they'll never forget.
After his relationship with the other animals improved over the past nine months, Surly Squirrel is now loved by Liberty Park's Urban wildlife community, and currently overseeing an all-you-can-eat buffet at Maury's Nut Shop. This worries Andie, as she prefers a more hard-working outlook on life. Despite Surly's reassurances all is well, Maury's Nut Shop is accidentally blown up by Mole after he forgets to cut down pressure from the boiler. Andie takes the opportunity to try getting the animals back to their roots by foraging for food in Liberty Park while Surly and Buddy try to find other food-packed places, but fail each time. Defeated, he and Buddy decide to return to the park.