Yogi and the gang visit Ranger Smith and put a show for a bus full of orphan children visiting Jellystone Park. Unfortunately, Wally Gator doesn't have any talents and has to be a curtain puller. An entertainer named Hilarious P. Prankster, as a revenge for being refused by Yogi to be in the show, tricks Wally by offering to make him a comedian. As a result, Wally starts playing mean pranks on everybody and starts losing his friends, until they learn Hilarious is behind Wally's new hobby.
Yogi gets homesick and the animals go back to Jellystone Park where there is a big party planned for him. However, the Gossipy Witch of the West (who works for Ranger Smith with various professions) finds and spreads rumors all over the park, including one that Yogi is engaged to Cindy and planning to leave the gang forever. At the same time, Chief Short (after seeing the ark and animals) is sure that a big flood is coming and investigates by dressing up as a giant chicken.
After a storm, Yogi and his gang stop on an island and need to get more supplies. They meet Mr. Waste (he actually introduces himself as "Mr. Plenty") who encourages the animals to stay on his island and use everything as much as possible. The animals are happy thinking they found a paradise, but Atom Ant (who as an ant doesn't believe in wasting things) soon finds what Mr. Plenty is really up to.
The gang stops at "Smog City," a city covered in smog by Mr. Smokestack Smog's smog factory where he has convinced everybody that smog is good for them. He encounters Magilla Gorilla, Peter Potamus and Snagglepuss (who are looking for bananas for the ark) and tries to convince them into his way of thinking.
Wally Gator is sad because he is the only one of the animals who doesn't own a surfboard or any other beach gear. However, a pirate named Captain Swashbuckle Swipe decides to teach him the "easy way" (stealing). Yogi enlists the help of Fumbo-Jumbo the Masked Avenger: a goofy Zorro-like elephant to help recover their stuff.
Cindy Bear gets ownership of a dude ranch, and all the animals come to help her get it ready for the guests. But a bad guy named Mr. Hothead (who goes under the name Mr. Coolhead) uses his invention to make all the animals except for Yogi, Cindy, and Boo Boo lose their tempers. Since they can't work together, the boys won't finish their jobs in time and Mr. Hothead will trick Cindy into selling him her ranch. Will they learn to work things out in a calm way without losing their tempers?
Broadcast in 1972 as part of "The ABC Saturday Superstar Movie" series, "Yogi's Ark Lark" was an animated Hanna-Barbera made-for-TV movie intended to raise ecological awareness. It also served as the pilot for the cartoon series "Yogi's Gang." This one-hour feature was later turned into a two-part series finale of "Yogi's Gang" with new intro and concluding sequences created for each episode. To fit the feature into these new time frames, some of its sequences were shortened or edited out. Aside from its environmental message, "Yogi's Ark Lark" marked a milestone in Hanna-Barbera's history: a wide range of the studio’s characters were united in one story for the first time. This set the tone for future series and specials, such as "Laff-a-Lympics" and "Yogi’s First Christmas."