As the girls, who are no longer able to transform, become curious as to why the Melody of Sorrow hasn't been sung yet, Otokichi announces the completion of his pipe organ, ready for a concert on Christmas Day. On the day of the concert however, Falsetto arrives to sing the Melody of Sorrow, with Otokichi's efforts of stopping it thwarted by Noise, who evolves into his true form, turns all of Kanon Town's inhabitants into stone and creates a Negatone from the Legendary Score to attack the girls before escaping to attack Major Land. As Otokichi strives to protect the girls, Hibiki notices the sorrow withing the Legendary Score and expresses her own feelings. As the others back her up, their feelings reach the Legendary Score, who returns their Heart Tone Keys, allowing them to transform and defeat the Negatone, returning the score to its original state. Using the Shirabe House as an airship, the Cures and Otokichi travel to Major Land to stop Noise.
ついに不幸のメロディの楽譜が完成してしまった。響たちはプリキュアに変身することもできず、どうすれば良いか思い悩む。そんな彼女たちに音吉が、パイプオルガンが完成したから心配はないと話す。