A specialist wildlife cameraman from the BBC's Planet Earth series arrives at the park to film Longleat's most impressive feeders. His subject is a chameleon who has a tongue which flies out four times faster than a jet fighter. But what if it doesn't want breakfast? Meanwhile, Kate Humble and Ben Fogle discover the lengths keepers go to for the animal orphans, while Jean Johansson sets up a tug of war for the lion pride. Elsewhere, the stakes couldn't be higher as vital testing for a killer disease continues. But when tragedy strikes, can the keepers recover from the unthinkable?