The film depicts Steve Coogan playing himself as an arrogant actor with low self-esteem and a complicated love life. Coogan is playing the eponymous role in an adaptation of ''The Life and Opinions of Tristram Shandy, Gentleman'' being filmed at a stately home. He constantly spars with actor Rob Brydon, who is playing Uncle Toby and believes his role to be of equal importance to Coogan's, calling himself the "co-lead".
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