A series in which Mater tells McQueen about his adventures in various fields.
The short film opens with Mater playing pranks on the other residents of Radiator Springs: moving Red's flowers, bursting out from a pile of tires to scare Guido and Luigi, dressing up as a cone vampire to scare Sally, attempting to scare Lizzie while she's sleeping (which is unsuccessful), moving Fillmore's gas cans back outside while he takes them in, and jumpscaring Lightning McQueen when he believes Mater is hiding behind a pile of cans ready to jump out (what was really behind the pile of cans is one of Mater's signs). This sequence culminates with Mater teasing McQueen as if he had seen "the Spooklight," a U.S. Route 66 legend recounted to Pixar by Dean Walker of the Route 66 association Kansas Historic Route 66 Association.
Lightning McQueen and Mater are driving in Radiator Springs and drive past Red, who is hosing some plants near Stanley. Mater tells Lightning that he used to be a firetruck, but Lightning doesn't believe Mater.
A routine tow lands Mater in Tokyo, where he is challenged to a drift-style race against a nefarious gang leader and his posse of ninjas. With the help of his friend, 'Dragon' Lightning McQueen, and some special modifications, Mater attempts to drift to victory and become Tow-ke-O Mater, King of all Drifters.
Mater finds a small UFO called Mator and they have a night out. Later, when Mator is captured by military forces, Mater sneaks up and saves him with the help of Lightning McQueen and the UFO's mother.
Mater and Lightning McQueen are ready to rock! When Mater's garage band, Mater and the Gas Caps, records a hit song, Mater becomes a rock legend. Then Lightning joins Mater's band for the rock concert of the century!
As a professional monster truck wrestler, Mater must work his way up through the ranks from an amateur tow truck to World Champion Monster Truck Wrestler. But rival wrestlers I-Screamer, Captain Collision, and The Rasta Carian aren't about to give up without a fight.
Mater is the first tow truck on the moon! When an auto-naut gets stranded on his lunar mission, it's up to Mater to venture into space and tow him back to Earth (with a little help from his friend Lightning McQueen, of course).
Detective Mater is hot on the trail of a dastardly car-napping! When Tia's sister goes missing, Mater is the only car she can turn to. With this tow truck on the case, anything can happen.
When a clock lands on Mater's engine, he travels back in time to 1909 where he meets Stanley, an ambitious young car on his way to California.